Theloderma khoii Ninh & Nguyen & Nguyen & Hoang & Siliyavong & Nguyen & Ziegler 2022, sp. nov.

Theloderma khoii sp. nov. urn:lsid:zoobank.org:act: 491C280C-F0CA-475E-8DC9-377603DCA558 Figs 2–3 Diagnosis Theloderma khoii sp. nov. shows the diagnostic characters of the genus Theloderma, for instance a distinct tympanum, round canthus rostralis, bony ridges from canthus rostralis to occiput absent, and skin of head not co-ossified to the skull, and also molecular analyses revealed it to be nested within Theloderma. The new species is distinguished from its congeners and other small rhacophorid species by a combination of the following characters: 1) large-sized frog of the genus Theloderma (SVL 52.2 mm in a single male, 59.4 mm in a single female); 2) head length and width equal; 3) vomerine teeth present; 4) snout pointed and truncated (SNL/SVL 16.3% in male; 16.0% in female); 5) eye large, eye diameter about a half of snout length (ED/SNL 54.60% in male; 58.9% in female), spinules on upper eyelid; 6) tibiotarsal articulation reaching to the posterior border of the eye or the tip of the snout; 7) dorsal skin very rough, with large irregular gland ridges and warts, ventral surface granular; 8) fingers rudimentary webbed, toes almost four-fifths webbed, tips of all digits dilated but all considerably smaller than tympanum; 9) dorsal surface mossy green or olive mottled with dark magenta; 10) lateral and ventral surfaces black with irregular lemon yellow pattern and some white granules; 11) male with large nuptial pads and an inner vocal sac. Etymology The specific epithet is in honor of Professor Dr Le Vu Khoi of the VNU Hanoi University of Science, Ha Noi, Vietnam, in recognition of his great support of teaching, mentoring, research and conservation work in Vietnam. As common names we suggest Khoi’s Mossy Frog (English) and Ếch cây sần khôi (Vietnamese). Type material Holotype VIETNAM • adult ♂; northern Vietnam, Ha Giang Province, Quan Ba District, in the karst forest near Tung Vai Commune; 1641 m a.s.l; 8 Jun. 2020; H.Q. Nguyen leg.; VNMN 012757. Paratype VIETNAM • adult ♀; same collection data as for ...