Alpheus edwardsii, sensu lato

Alpheus edwardsii (Audouin, 1826) sensu lato (Fig. 2) Athanas Edwarsii Audouin 1826: 91 (lap. cal.). Athanas Edwardsii. — Audouin 1827: 274. Alpheus Edwardsii. — Coutière 1898: 249, figs. 4, 5; Nobili 1901: 2; Banner & Banner 1973a: 1141, fig. 1; Kazmi & Kazmi 1979: 154; Banner & Banner 1981: 22, 92; Banner & Banner 1982: 270, fig. 83; Lewinsohn & Galil 1982: 208; Afzal et al. 1986: 338; De Grave & Ashelby 2011: 3; Naderloo & Türkay 2012: 9; Naderloo et al. 2013: 448; Dehghani et al. 2019a: 484, fig. 3A, B. Alpheus edwardsii sensu lato.— Anker & De Grave 2016: 352, figs. 10, 11. (?) Alpheus Audouini Coutière 1905: 911, pl. 87, fig. 52. Not Alpheus edwardsii. — Al-Maliky et al. 2017: 49, figs. 2–4 [= Alpheus lobidens De Haan, 1849]. Material examined. Kuwait: 1 male, OUMNH. ZC. 2020.01.018, Kubbar Island, 29°04’18.93”N 48°29’18.74”E, depth 2–3 m, sand flat with sparse rubble, under coral rocks, leg. S. De Grave, 26.ix.2019 [fcn KUW-017]; 2 males, 1 female, KISR, Miscan Island, sandy-rocky intertidal, leg. M. Al-Kandari et al., 15.xii.2019. General distribution. Species complex (see Remarks below), reported from throughout the Indo-West Pacific, from the Red Sea, Arabian Gulf (Iran, Kuwait, UAE) and South Africa across the Indian Ocean to Japan, Australia and French Polynesia (e.g., Banner & Banner 1973a, 1981, 1982, 1983; Chace 1988; Naderloo & Türkay 2012; Anker & De Grave 2016; Dehghani et al. 2019a; present study); Lessepsian migrant into the eastern Mediterranean Sea (Forest & Guinot 1958; Holthuis & Gottlieb 1958). Ecology. Variable (species complex, see below); found both in the intertidal and shallow subtidal (to at least 36 m), rocky and mixed rocky-sandy shores, also on coral reefs and associated coral rubble habitats (reef flats); typically, under large rocks and coral rubble (e.g. Banner & Banner 1982; Corfield & Alexander 1995; present study). Remarks. Without a comprehensive revision of the A. edwardsii species complex (cf. Anker & De ...